The study, "Migratory Appendicular muscles precursor cells in the common ancestor to all vertebrates," published in Nature Ecology & Evolution, was a collaboration between several centers in Japan and Spain, namely the Tokyo Tech, RIKEN Center for Life Science Technologies, the University of Tokyo as well as the Center for Genomic Regulation and Universitat Pompeu Fabra. Using catshark embryos as their research model, the investigators confirmed that Lbx1-positive cells are found in cartilaginous fish fin as well as in hypobranchial (pertaining to the segment below the gills) muscles, and that these are formed via the mechanism that has been established in land dwelling vertebrates as well as in bony fish.
In addition to Lbx1, Mikiko Tanaka at Tokyo Tech and team found other genes that are involved in fin muscle formation...
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